张敬，男，1989年2月生，副教授，硕士生导师。攻读博士学位期间赴美国罗格斯大学公派留学2年，并获中国草学会“王栋奖学金”、南京农业大学“优秀博士学位论文”等荣誉。主要从事黑麦草耐逆、产量和品质等性状调控机理与种质创新研究。主持国家自然科学基金重点项目（子课题）、面上项目（2项）、国家自然科学青年基金、江苏省青年基金、江苏省农业科技自主创新基金和中国博士后基金（2项）项目。相关研究结果以第一或通讯（含共同）作者在The Plant Journal（2篇, 2025; 2026）, Plant Physiology（2025）, Horticulture Research（2篇, 2026a; 2026b）, Plant, Cell & Environment（2022）和Journal of Experimental Botany （2016）等国际知名期刊上发表学术论文20余篇，其中ESI高被引论文2篇。培育‘南黑4号’（1/4）和‘南农6号’（2/5）黑麦草新品系（分别于2024年和2025年进入国家新品种区域试验）。

一、教育经历

  2013.09-2017.06 南京农业大学，草学，博士 

  2014.09-2016.09 美国罗格斯大学，植物学系，联合培养博士

  2011.09-2013.06 南京农业大学、江苏省农科院，草业科学，硕士

  2007.09-2011.06 安徽农业大学，生物科学类，理学学士

二、工作经历

  2023.01-至今南京农业大学，草业学院，副教授

  2020.02-2022.12, 南京农业大学，草业学院，讲师

  2017.09-2020.01, 南京农业大学，草业学院，师资博士后

三、主持科研项目

1. 张敬：柠檬酸合酶LpCSY调控多年生黑麦草耐热性的机制研究，国家自然科学基金面上项目（32674138），2027.01-2030.12，主持，50万。

2. 张敬：蔗糖-果聚糖代谢关键酶1-SST乙酰化修饰调控多年生黑麦草耐热性的机制研究，国家自然科学基金面上项目（32371778），2024.01-2027.12，主持，50万。

3. 张敬：西南黑麦草耐热关键基因挖掘及分子调控机制解析，国家自然科学基金重点项目（U23A20218），2024.01-2027.12，子课题负责人（子课题59万）。

4. 张敬：叶绿素b合成基因LpCAO响应遮荫并调控多年生黑麦草耐阴性的机制研究，中国博士后基金面上项目，2023.09-2025.09，主持，8万。

5. 张敬：细胞分裂素转录因子LpARR11调控多年生黑麦草衰老及叶绿素降解的分子机制，国家自然科学青年基金（31802117），2019.01-2021.12，主持，26万。

6. 张敬：多年生黑麦草耐热种质筛选及新品系创制，江苏省农业科技自主创新资金项目[CX(21)3004]，2021.07-2023.08，主持，20万。

7. 张敬：LpARR11介导细胞分裂素调控多年生黑麦草叶片衰老的分子机制，江苏省自然科学青年基金（BK20180546），2018.07-2021.06，主持，20万。

8. 张敬：多年生黑麦草种质资源耐热性评价及相关分子标记的发掘，中国博士后基金面上项目（2017M621764），2018.07-2020.06，主持，5万。

四、学术论文

第一作者或通讯作者论文（#代表共一作者；*代表通讯作者）

1. Yin T, Zhang Z, Hao H, Yang Y, Wang L, Zhou Q, Zhang A, Chi Y, Zhang J*, Xu B*. A bHLH transcription factor, LpCbDR2, activates LpNOL to regulate senescence, heat tolerance, and crude protein content in perennial ryegrass. The Plant Journal. 2026, 126, e71007.

2. Yin T, Guo H, Zhu Y, Yang Y, Hao H, Liu X, Lou J, Xie C, Wang Y, Yan H, Huang L, Li Y, Yan S, Chi Y, Xu B*, Zhang J*. Chromosome-level genome assembly of Origanum vulgare subsp. hirtum reveals evolutionary insights and regulatory modules in terpenoid biosynthesis. Horticulture Research. 2026, 13(7), uhag093.

3. Hao H, Zhou Q, Yin T, Dong C, Zhang Z, Chi Y, Zhang J*, Xu B*. LpCbDR1 regulates leaf senescence and drought tolerance in perennial ryegrass by activating the chlorophyll b reductase gene and stress-related genes. Horticulture Research. 2026, 13(5), uhag030.

4. Yang Y, Xing J, Hao H, Guan H, Yang H, Yin T, Chi Y, Xu B*, Zhang J*. LpSAPK9 phosphorylates and activates LpABF2/LpABF3 to transactivate expression of chlorophyll catabolic genes and promote leaf senescence in perennial ryegrass. The Plant Journal, 2025, 123, e70477.

5. Yang H, Yao J, Hao H, Jiang L, Li J, Chi Y, Zhang J*, Xu B*. Promoter editing of LpNOL generates stay-green perennial ryegrass with improved forage quality and heat tolerance. Plant Physiology, 2025, 199, kiaf402.

6. Zhang J, Li H, Huang X, Xing J, Yao J, Yin T, Jiang J, Wang Pu, Xu B*. STAYGREEN-mediated chlorophyll a catabolism is critical for photosystem stability during heat-induced leaf senescence in perennial ryegrass. Plant, Cell & Environment, 2022, 45(5):1412-1427. ESI高被引论文.

7. Zhang J, Yu G, Wen W, Ma X, Xu B*, Huang B*. Functional characterization and hormonal regulation of the pheophytinase gene LpPPH controlling leaf senescence in perennial ryegrass. Journal of Experimental Botany, 2016, 67(3), 935-945.

8. Guan H, Xie C, Jia W, Ouyang R, Xia Y, Fu Y, Man Y, Hou H, Chi Y, Xu B*, Zhang J*. Acetic acid suppresses perennial ryegrass growth in vitro by disrupting gibberellin and auxin homeostasis and restricting cell expansion. Journal of Plant Growth Regulation. 2026,1-12 (https://doi.org/10.1007/s00344-026-12310-z).

9. Yang H, Zhang Q, Li Z, Zhao R, Liu X, Chi Y, Zhang J*, Xu B. Molecular characterization of the cytokinin responsive type-B response regulators in perennial ryegrass highlights LpARR10 conferring salt tolerance via trans-activating SOS1 and SOS3. Environmental and Experimental Botany. 2024, (217):105566.

10. Xing J, Yang Y, Zhang Q, Yin T, Zhao R, Guan H, Liu X, Chi Y, Zhang J*. A cytochrome P450 gene, LpCYP72A15, confers drought tolerance in perennial ryegrass. Grass and Forage Science. 2023, 12, 05.

11. Zhang J*, Chen Y, Chai M, Shabala S, Wang K, Zhang J.L*. Editorial: Adaptation mechanisms of grass and forage plants to stressful environments, volume Ⅱ. Frontiers in Plant Science. 2023, 14:1323841.

12. Zhang J, Chai M, Shabala S, Wang K, Zhang J.L*. Editorial: Adaptation mechanisms of grass and forage plants to stressful environments. Frontiers in Plant Science. 2023, 14:1132198.

13. Song G#, Zhang J#, Wang Y, Ji Y, Fang Z, Cai Q*, Xu B*. Overexpression of PvBiP2 improved biomass yield and cadmium tolerance in switchgrass (Panicum virgatum L.). Journal of Hazardous Materials, 2023, 446:130648.

14. Xing J, Zhao R, Zhang Q, Huang X, Yin T, Zhang J*, Xu B*. Genome-wide identification and characterization of the LpSAPK family genes in perennial ryegrass highlight LpSAPK9 as an active regulator of drought stress. Frontiers in Plant Science, 2022, 13:922564.

15. Zhang J, Zhang Q, Xing J, Li H, Miao J, Xu B*. Acetic acid mitigated salt stress by alleviating ionic and oxidative damages and regulating hormone metabolism in perennial ryegrass (Lolium perenne L.). Grass Research, 2021, 1: 3.

16. Zhang J, Wen W, Li H, Lu Q, Xu B*, Huang B. Overexpression of an aquaporin gene PvPIP2;9 improved biomass yield, protein content, drought tolerance and water use efficiency in switchgrass (Panicum virgatum L.) Global Change Biology Bioenergy, 2020, 00:1-13.

17. Zhang J, Li H, Jiang Y, Li H, Zhang Z, Xu Z, Xu B*, Bingru Huang. Natural variation of physiological traits, molecular markers, and chlorophyll catabolic genes associated with heat tolerance in perennial ryegrass accessions. BMC Plant Biology, 2020, 20:520.

18. Zhang J, Xing J, Lu Q, Yu G, Xu Bin *, Huang Bingru*. Transcriptional regulation of chlorophyll-catabolic genes associated with exogenous chemical effects and genotypic variations in heat-induced leaf senescence for perennial ryegrass. Environmental and Experimental Botany, 2019, 11.

19. Xu B, Yu G, Li H, Xie Z, Wen W, Zhang J*, Huang B*. Knockdown of STAYGREEN in perennial ryegrass (Lolium perenne L.) leads to transcriptomic alterations related to suppressed leaf senescence and improved forage quality. Plant & Cell Physiology, 2019, 60, 202-212.

20. Xu B, Li H, Li Y, Yu G, Zhang J*, Huang B*. Characterization and transcriptional regulation of chlorophyll b reductase gene NON-YELLOW COLORING 1 associated with leaf senescence in perennial ryegrass (Lolium perenne L.). Environmental and Experimental Botany, 2018, 149.

21. Shi Y#, Zhang J#, Li, Li M, Huang B*. Butanediol-enhanced heat tolerance in Agrostis stolonifera in association with alteration in stress-related gene expression and metabolic profiles. Environmental and Experimental Botany, 2018, 153:209-217.

22. Zhang J, Shi Y, Zhang X, Du H, Xu B*, Huang B*. Melatonin suppression of heat-induced leaf senescence involves changes in abscisic acid and cytokinin biosynthesis and signaling pathways in perennial ryegrass (Lolium perenne L.). Environmental and Experimental Botany, 2017, 138:36-45. ESI高被引论文.

23. Zhang J, Li H, Xu B, Li J, Huang B*. Exogenous melatonin suppresses dark-induced leaf senescence by activating the superoxide dismutase-catalase antioxidant pathway and down-regulating chlorophyll degradation in excised leaves of perennial ryegrass (Lolium perenne L.). Frontiers in Plant Science, 2016, 7.

24. Jespersen D#, Zhang J#, Huang B*. Chlorophyll loss associated with heat-induced senescence in bentgrass. Plant Science, 2016, 249,1-12.

25. 关昊（本科生），姚佳明，张欣栎，王硕，宋欣瑶，刘信宝，张敬*，徐彬. 多年生黑麦草LpPIL15-like基因转录模式及耐逆功能分析. 中国草地学报, 2024, 46(2), 24-34.

26. 张晴，邢静，姚佳明，殷庭超，黄心如，何悦，张敬*，徐彬. 多年生黑麦草细胞分裂素信号通路B类ARR转录因子LpARR10的耐镉功能分析. 草业学报, 2022, 31(5),135-143.

27. 姚佳明，何悦，郝欢欢，黄心如，张敬*，徐彬. 多年生黑麦草LpPIL5基因特征分析及转录调控. 草业学报, 2022, 31(9), 1-13.

五、荣誉及奖项

中国草学会王栋奖学金（2015.10，张敬）

南京农业大学优秀博士学位论文（2018.12，张敬）

全国草学类本科专业2025年青年教师授课比赛一等奖（2025.09，张敬）

南京农业大学校级优秀毕业论文一等奖2项（2024和2026，导师，张敬）

南京农业大学2024届研究生校长奖学金（2024.06，合作指导老师，张敬）

国家留学基金委公派留学（2014.09-2016.09，张敬）

全国高校农林类专业微课教学比赛—教学风采奖（2022.11，张敬）